This study was conducted to determine the possible number of developmental modules defining the shape of the hind wing of coconut hispid beetle B. longissima. Geometric Morphometric analysis (GM) was used to describe the shape of the wings summarized via Procrustes analysis. A total of 180 landmark points trace and outline the margins of the wings as well as the major veins. Modularity and Integration Analysis (MINT) software was used as a tool to test a priori models of variational modularity in multidimensional data. Thirteen a priori models of variational modularity in the GM hind wing data were tested using the γ* (Gamma*) test for goodness of fit (GoF) statistics by comparing the observed and expected covariance matrices. Results showed that both male and female coconut hispid beetle hind wing is organized into four modules, the partition of cubitus and media, radius and media, costa+subcosta and radius and cubitus and anal vein. Differences in ranks of the top 3 models were observed in male and female left wing and right wing. It was hypothesized to be caused by effects associated with the signalling sources for position and morphology of pattern elements which only covers short distances.
